1. Use room-temperature butter eggs and buttermilk for best texture 2. avoid overmixing after adding flour to keep cupcakes tender 3. substitute plain kefir or milk plus 1 tbsp lemon juice for buttermilk 4. for extra lemon flavor brush cooled cupcakes with a light lemon syrup before frosting 5. store airtight at room temperature up to 1 day or refrigerate up to 4 days and bring to room temperature before serving